Mark Webster

Balaguera-Reina, S. A., M. Angulo-Bedoya, J. F. Moncada-Jimenez, M. Webster, I. J. Roberto, and F. J. Mazzotti. 2022. Update: Assessing the evolutionary trajectory of the Apaporis caiman (Caiman crocodilus apaporiensis, Medem 1955) via mitochrondrial molecular markers. Biological Journal of the Linnean Society.

Manzuk, R. A., A. C. Maloof, J. A. Kaandorp, and M. Webster. 2022. Braching archaeocyathids as ecosystem engineers during the Cambrian radiation. Geobiology. [Link to PDF]

Sundberg, F. A., M. Webster, and G. Geyer. 2022. Biostratigraphical significance of a new trilobite fauna from the Harkless Formation (upper Stage 4, Series 2, Cambrian), Nevada, USA. Lethaia 55 (3): 1-12.

Sundberg, F. A., and M. Webster. 2022. 'Ptychoparioid' trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada. Journal of Paleontology 96 (4): 886-920. [Link to PDF]

Sundberg, F. A., and M. Webster. 2021. Corynexochine trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada. Journal of Paleontology 95 (6): 1241-1258. [Link to PDF]

Hughes, N. C., J. M. Adrain, J. D. Holmes, P. S. Hong, M. J. Hopkins, J.-b. Hou, A. Minelli, J. Peng, T.-Y. Park, J. R. Paterson, J. Peng, M. Webster, X.-G. Zhang, X.-L. Zhang, and G. Fusco. 2021. Articulated trilobite ontogeny: suggestions for a methodological standard. Journal of Paleontology 95 (2): 298-304. [Link to PDF]

Moore, J. L., S. M. Porter, M. Webster, and A. C. Maloof. 2021. Chancelloriid sclerites from the Dyeran-Delamaran ('Lower-Middle' Cambrian) boundary interval of the Pioche-Caliente region, Nevada, USA. Papers in Palaeontology. 7 (1): 565-623. doi:10.1002/spp2/1274.

Webster, M., and F. A. Sundberg. 2020. Nature and significance of intraspecific variation in the early Cambrian oryctocephalid trilobite Oryctocephalites palmeri Sundberg and McCollum, 1997. Journal of Paleontology 94 (1): 70-98. [Link to PDF]

Webster, M. 2019. Morphological homeostasis in the fossil record. Seminars in Cell and Developmental Biology 88: 91-104. [Link to PDF]

Webster, M., and S. J. Hageman. 2018. Buenellus chilhoweensis n. sp. from the Murray Shale (lower Cambrian Chilhowee Group) of Tennessee, the oldest known trilobite from the Iapetan margin of Laurentia. Journal of Paleontology 92 (3): 442-458. [Link to PDF]

Landing, E., and M. Webster. 2018. Iapetan rift-passive margin transition in NE Laurentia and eustatic control on continental slope oxygenation, Taconic slate colors, and Early Paleozoic climate. In Grover, T. W. (ed.), Guidebook to Field Trips in New York, Vermont, and Massachusetts. 110th New England Intercoll. Geol. Conf. and 90th New York State Geol. Assoc. Meeting, Lake George, NY, October 12-14. [PDF Link]

Webster, M., and E. Landing. 2016. Geological context, biostratigraphy and systematic revision of late early Cambrian olenelloid trilobites from the Parker and Monkton formations, northwestern Vermont, U.S.A. Australasian Palaeontological Memoirs 49: 193-240. [PDF Link]

Webster, M. 2015. Ontogeny and intraspecific variation of the early Cambrian trilobite Olenellus gilberti, with implications for olenelline phylogeny and macroevolutionary trends in phenotypic canalization. Journal of Systematic Palaeontology 13 (1): 1-74. (Published online in 2014) [Link to PDF]

Webster, M., and L. L. Bohach. 2014. Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia. Zootaxa 3824 (1): 1-66. [Link to PDF]

Webster, M., and M. L. Zelditch. 2011. Evolutionary lability of integration in Cambrian ptychoparioid trilobites. Evolutionary Biology 38: 144-162. [PDF Link][Link to R code]

Webster, M., and M. L. Zelditch. 2011. Modularity of a Cambrian ptychoparioid trilobite cranidium. Evolution & Development 13 (1): 96-109. [PDF Link]

Webster, M. 2011. The structure of cranidial shape variation in three early ptychoparioid trilobite species from the Dyeran-Delamaran (traditional "Lower-Middle" Cambrian) boundary interval of Nevada, U. S. A. Journal of Paleontology 85 (2): 179-225. [Request a PDF]

Webster, M. 2011. Upper Dyeran trilobite biostratigraphy and sequence stratigraphy in the southern Great Basin. Pp. 121-154 in J. S. Hollingsworth, F. A. Sundberg, and J. R. Foster (eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin 67. Flagstaff, Arizona. [PDF Link]

Webster, M. 2011. Litho- and biostratigraphy of the Dyeran-Delamaran boundary interval at Frenchman Mountain, Nevada. Pp. 195-203 in J. S. Hollingsworth, F. A. Sundberg, and J. R. Foster (eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin 67. Flagstaff, Arizona. [PDF Link]

Webster, M. 2011. Litho- and biostratigraphy of the Dyeran-Delamaran boundary interval in the Pioche-Caliente region, Nevada. Pp. 203-215 in J. S. Hollingsworth, F. A. Sundberg, and J. R. Foster (eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin 67. Flagstaff, Arizona. [PDF Link]

Webster, M., L. B. McCollum, and F. A. Sundberg. 2011. Upper Dyeran and Lower Delamaran litho- and biostratigraphy of the northern Groom Range, Nevada. Pp. 226-236 in J. S. Hollingsworth, F. A. Sundberg, and J. R. Foster (eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin 67. Flagstaff, Arizona. [PDF Link]

Webster, M. 2011. Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. Pp. 236-246 in J. S. Hollingsworth, F. A. Sundberg, and J. R. Foster (eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin 67. Flagstaff, Arizona. [PDF Link]

Webster, M., and H. D. Sheets. 2010. A practical introduction to landmark-based geometric morphometrics. Pp. 163-188 in J. Alroy and G. Hunt (eds.), Quantitative Methods in Paleobiology. Paleontological Society Papers, Volume 16. [PDF Link]

Webster, M. 2009. Systematic revision of the Cambrian trilobite Bathynotus Hall, 1860, with documentation of new occurrences in western Laurentia and implications for intercontinental biostratigraphic correlation. Memoirs of the Association of Australasian Paleontologists 37: 369-406. [PDF Link]

Hopkins, M. J., and M. Webster. 2009. Ontogeny and geographic variation of a new species of the corynexochine trilobite Zacanthopsis (Dyeran, Cambrian). Journal of Paleontology 83 (4): 524-547. [Request a PDF]

Webster, M. 2009. Ontogeny, systematics, and evolution of the effaced early Cambrian trilobites Peachella Walcott, 1910 and Eopeachella new genus (Olenelloidea). Journal of Paleontology 83 (2): 197-218. [Request a PDF]

Webster, M., and M. L. Zeldtich. 2008. Integration and regulation of developmental systems in trilobites. Pp. 427-433 in Rábano, I., Gozalo, R., and García-Bellido, D (eds.), Advances In Trilobite Research. Cuadernos del Museo Geominero 9. Instituto Geológico y Minero de España, Madrid. [PDF Link]

Hopkins, M. J., and M. Webster. 2008. Morphological and ontogenetic change in the "Early" Cambrian trilobite Zacanthopsis during an interval of environmental change. Pp. 185-187 in Rábano, I., Gozalo, R., and García-Bellido, D (eds.), Advances In Trilobite Research. Cuadernos del Museo Geominero 9. Instituto Geológico y Minero de España, Madrid. [PDF Link]

Webster, M., R. R. Gaines, and N. C. Hughes. 2008. Microstratigraphy, trilobite biostratinomy, and depositional environment of the "Lower Cambrian" Ruin Wash Lagerstätte, Pioche Formation, Nevada. Palaeogeography, Palaeoclimatology, Palaeoecology 264: 100-122. [Request a PDF]

Webster, M. 2007. Paranephrolenellus, a new genus of Early Cambrian olenelloid trilobite. Memoirs of the Association of Australasian Paleontologists 34: 31-59. [PDF Link]

Webster, M. 2007. Ontogeny and evolution of the Early Cambrian trilobite genus Nephrolenellus (Olenelloidea). Journal of Paleontology 81 (6): 1168-1193. [PDF Link]

Webster, M. 2007. A Cambrian peak in morphological variation within trilobite species. Science 317: 499-502. [PDF Link]

Webster, M., and M. L. Zelditch. 2005. Evolutionary modifications of ontogeny: heterochrony and beyond. Paleobiology 31 (3): 354-372. [PDF Link]

Fusco, G., N. C. Hughes, M. Webster, and A. Minelli. 2004. Exploring developmental modes in a fossil arthropod: growth and trunk segmentation of the trilobite Aulacopleura konincki. American Naturalist 163 (2): 167-183.

Webster, M., P. M. Sadler, M. A. Kooser, and E. Fowler. 2003. Combining stratigraphic sections and museum collections to increase biostratigraphic resolution: Application to Lower Cambrian trilobites from southern California. Pp. 95-128 in P. J. Harries (ed.), High-Resolution Approaches in Stratigraphic Paleontology. Topics in Geobiology, Volume 21. Kluwer Academic Publishers, Dordrecht.

Webster, M., H. D. Sheets, and N. C. Hughes. 2001. Allometric patterning in trilobite ontogeny: Testing for heterochrony in Nephrolenellus. Pp. 105-144 in M.L. Zelditch (ed.), Beyond Heterochrony: The Evolution of Development. John Wiley and Sons, New York.

Dunlop, J. and M. Webster. 1999. Fossil evidence, terrestrialization and arachnid phylogeny. Journal of Arachnology 27: 86-93.

Webster, M., and N. C. Hughes. 1999. Compaction-related deformation in Cambrian olenelloid trilobites and its implications for fossil morphometry. Journal of Paleontology 73: 355-371.

Bowden, A., M. Webster, and T. Mitcham. 1997. Salthill Quarry Geology Trail. Geologists' Association Guide No. 58. Geologists' Association, London. 30 pp.